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Rye is a town and civil parish in the Rother district of East Sussex, England, two miles (three kilometres) from the sea at the confluence of three rivers: the Rother, the Tillingham and the Brede. An important member of the mediaeval Cinque Ports confederation, it was at the head of an embayment of the English Channel, and almost entirely surrounded by the sea.

At the 2011 census, Rye had a population of 4,773. Its historical association with the sea has included providing ships for the service of the Crown in time of war, and being involved in smuggling. The notorious Hawkhurst Gang used its ancient inns The Mermaid Inn and The Olde Bell Inn, which are said to be connected to each other by a secret passageway.

Those historic roots and its charm make it a tourist destination, with hotels, guest houses, B&Bs, tea rooms, and restaurants. Rye has a small fishing fleet, and Rye Harbour has facilities for yachts and other vessels.

The name of Rye is believed to come from the West Saxon ieg meaning island. Medieval maps show that Rye was originally located on a huge embayment of the English Channel called the Rye Camber, which provided a safe anchorage and harbour. Probably as early as Roman times, Rye was important as a place of shipment and storage of iron from the Wealden iron industry.

Rye, as part of the Saxon Manor of Rameslie, was given to the Benedictine Abbey of Fécamp in Normandy by King Æthelred; it was to remain in Norman hands until 1247. The town of Rye is recorded in the 1086 Domesday Book as possessing 189 households, marking it as a significant settlement at the time. The cellars of the Mermaid Inn date from 1156.

As one of the two "Antient Townes" (Winchelsea being the other), Rye was to become a limb of the Cinque Ports. Confederation by 1189, and subsequently a full member. The protection of the town as one of the Cinque Ports was very important, due to the commerce that trading brought. One of the oldest buildings in Rye is Ypres Tower, which was built in 1249 as "Baddings Tower", to defend the town from the French, and was later named after its owner, John de Ypres. It is now part of the Rye Museum.

Rye received its charter from King Edward I in 1289, and acquired privileges and tax exemptions in return for ship-service for the crown. The "Landgate" (the only surviving one of four original fortified entrances to Rye) dates from 1329 in the early years of the reign of King Edward III. It is suitable only for light vehicles. In 2015, some 25 tonnes of pigeon excrement that had built up had to be removed from Landgate Arch for fear of damaging the ancient structure.

The River Rother originally took an easterly course to flow into the sea near what is now New Romney. However, the violent storms in the 13th century (particularly in 1250 and 1287) cut the town off from the sea, destroyed Old Winchelsea, and changed the course of the Rother. Then the sea and the river combined in about 1375 to destroy the eastern part of the town and ships began use the current area (the Strand) to unload their cargoes. Two years later, the town was sacked and burnt by the French, and it was ordered that the town walls be completed, as a defence against foreign raiders.

The South Gate at Rye, 1785, by Samuel Hieronymus Grimm
Rye was considered one of the finest of the Cinque Ports, though constant work had to be done to stop the gradual silting up of the river and the harbour. Also, a conflict arose between the maritime interests and the landowners, who gradually "inned" or reclaimed land from the sea on Romney and Walland Marsh, and thus reduced the tidal flows that were supposed to keep the harbour free of silt. Acts of Parliament had to be passed to enable the Rother to be kept navigable at all.

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